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That's a new stock 4R100 transmission, standard equipment on the F-150 Lightning. Actually, it's not quite the same as the tranny you'd find in a regular 5.4 F-150. This gearbox has the heavier-duty diesel transmission guts stuffed into the gasoline engine transmission case. It's an expensive and bulletproof transmission designed to handle the plentiful torque pumped out by the blown 5.4L engine. The custom transmission crossmember is fastened to the newly added subframe connectors that run the length of the floorpan and even extend up into the passenger compartment.